Jake received his BS in Zoology from Washington State University. As an undergraduate, he worked with Dr. William Snyder studying biodiversity in ecosystem functioning, with a focus on insect predator diversity. After graduating, Jake worked with Dr. Harmit Malik at the Fred Hutchinson Cancer Research Center in Seattle, WA. There, Jake studied the molecular evolution of retroelement and viral restriction factors in primates with an emphasis on how pathogens have shaped the evolution of host restriction factors like APOBEC3A. In the Hoekstra lab, Jake is studying the function, evolution and genetics of whisker biology in wild mice.